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The questions "PUPscan" won't answer. Part 1

2/14/2017

 
By Carol Beuchat PhD
I was asked for my opinion about a new venture in the United Kingdom called "PUPscan" (Puppy Ultrasound Project), which has launched a Facebook page with the banner "Working towards a better future for our dogs". I'm all for anything that could improve the lives of our dogs, so I spent a little time checking it out.
First, a bit of what they say about themselves.

​On their new Facebook page, they state that they are using ultrasound to evaluate the hips of 6-8 week old puppies. They say that "the EBV method is not yet able to demonstrate genetic abnormalities" and it relies "on the present hip scoring system which does not give a diagnosis of genetic information." They note that "there is NO identified gene for what is described as 'Dysplasia, a term taken from the Greek to mean "bad" or abnormal hip. It is not a full diagnosis." They point out that hip dysplasia diagnosed at 2 years old could be the result of "trauma, infection, a fluid bild up in the hip or simply over weight or over exercised none of which are genetic." Further, they claim that "X-raying dogs at full skeletal maturity for hip evaluation over the past 30+ years has not reduced the prevalence of 'Dysplasia' confirming that that this label is not fit for purpose." Finally, they point out that "too many dogs are being removed from the gene pool based on a score that does not give us a genetic diagnosis."
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They claim to be offering a solution to these issues.
From their Facebook page:

With PUPscan any hip abnormalities are identified at 6-8 weeks of age. Screening at 6-8 weeks gives us the opportunity to identify the truly genetic abnormalities. With PUPscan we can screen for hip 'Dysplasia' in 6-8 week old puppies. Our Ultrasound Imaging needs no anaesthetic and is achieved in the natural load bearing position. Congenital genetic 'Dysplasia' can be identified, thus preventing many at risk breeds being excluded from your breeding program by being labelled 'Dysplastic' although with no proven genetic abnormality.
​

There are additional details under Frequently Asked Questions:
With Ultrasound diagnostics, we can distinguish between congenital and acquired hip and elbow problems in young puppies. Congenital means that you are born with it, but not everything you are born with is genetic...Congenital may be acquired while in the womb or during birth for example. We can obtain a diagnostic image within two or three minutes per joint. Entering the data on to a computer system together with the measurements that represent the development of the joint takes a further five minutes.

From a reprint of a published article (posted on the Facebook page) about them written by one of the people involved in the venture:
With PUPscan we can screen for hip problems in six week old puppies by ultra sound imaging which will identify hip problems in vulnerable breeds, thereby helping to avoid the exclusion of many dogs from Breeding Programmes that are labelled as "Dysplastic" but may not be genetically so...Most of these puppies may actually be born with normal hips...With PUPscan, we can scan puppies at 6 weeks, which can go to their new homes with a clear certificate if no abnormality is found.

The bottom line seems to be that PUPscan is arguing that by screening puppies at 8 weeks, they can distinguish between "genetic" dysplasia and hip abnormalities with non-genetic (i.e., environmental) causes.
​
My questions

On the basis of the information they provided, I submitted some basic questions to their facebook page:

  • What "measurements" are taken?
  • What evidence is there that whatever you are scoring is predictive of the development of hip dysplasia?
  • What value is there in imaging a cartilagenous joint that is yet to be formed?
  • What is the basis for your claim to be assessing "congenital genetic dysplasia"?​​
These seem like straightforward questions for which PUPscan should have been able to provide responses. But they didn't. I asked the same set of questions again and again, and they responded with non-answers. Finally, they said "I think that your inability to grasp the simplicity of our project which can only enhance your EBV's is disappointing because you need more than D.N.A. tests to breed healthy puppies."
This is my response.
No, I don't get the "simplicity" of your project, because you haven't offered any evidence that it has anything useful to offer. And you haven't answered the questions that would provide this information.
 
Here's the issue. The hip joints of puppies at 8 weeks are cartilage. There will be no evidence of hip dysplasia, genetic or otherwise, in an 8 week puppy, or a 4 week puppy, or a 2 week puppy, or any puppy with joints made of cartilage, unless it's been tossed off the roof a few times. So you can collect your money, hand out a certificate stating that you found no evidence of hip dysplasia at 8 weeks, and that would be true.
 
When a puppy goes on to develop dysplasia, you can then claim that it must be due to non-genetic factors, because the hips were "normal" at 8 weeks when you would have detected "genetic", "congenital" hip dysplasia. Of course, you do not assess laxity, which might have been at least nominally informative (e.g., as in a hip with broken ligament that can be subluxated). Riser would agree that it is biomechanics that results in the deformation of the hips that is manifested as hip dysplasia, but he ALSO makes it clear that genetic differences in body structure are also involved.
 
You know, it is important that we get things like this right. There are many breeders that will use the tools we provide them and will make every effort to do what is necessary to produce healthy, happy puppies. I would love to know that you have something useful to offer them. But I don't see it, and you're not offering the information that will convince me. Why? It doesn't escape me that you call this a "Project" and not a "test" or a "diagnosis" or some other term that denotes more than an activity of some sort. I can't help but be suspicious. You will be collecting money from people, but what useful information are you offering, and what is the PROOF that it is useful? This evidence should be front and center, yet you refuse to provide it. I'm not trying to be adversarial here, but do understand that ANYBODY using your "project" should be asking these questions. Surely you have answers?
 
While your certification that you find no evidence of hip dysplasia in the ultrasound of somebody's 8 week puppy might be true, it is, as far as I can tell, misleading and meaningless. I would argue further that you have done enough homework to know that the evaluations as described are specious and unethical.
 
I too am disappointed that I am still unable to grasp the simplicity of your "Project". If it was so simple, I would have thought it would be easy enough to explain. Plus, I still have my list of unanswered questions, which you have been given every opportunity to address. I'm still open to evaluating any information you might submit to the contrary before I write my review.

I did not receive additional information from PUPscan. However, they posted this response: 
Hi Carol, We in the United Kingdom find it impossible to explain why you in Califiornia are so hostile to an innovation that could improve the welfare of our dogs. Our Project compliments D.N.A analysis which is why our KC have asked us to collect samples from every puppy we scan for analyis and to include on their data base. Also where is your evidence that 50 years of hip scoring has eliminated 'Dysplasia'?
The next day, all of my comments and questions had been removed from the PUPscan Facebook page and I had been blocked. (Fortunately I had already saved a pdf of the page, anticipating that this might happen; you can download a copy below.)

​They had also added this:
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They also had this post on the Facebook page:
"I would like to remind people that this page is not here to promote other individual businesses; It is about information that we make available to all. FREE. If anyone feels they want to attend one of our seminar they are very welcome. PUPscan will not under any circumstances accept or totorate any attempts to discredit our work that is NOT evidence based. The KC are working with us to acheive a better understanding of canine health and welbeing. This is about working co-operativly in a multidisaplinary team."
An article about PUPscan published in Our Dogs (with a copy posted to their Facebook page) similarly mentions that the Kennel Club "had expressed their support for the scheme".
The Kennel Club was asked to comment and they provided this, a joint statement from the Kennel Club and The British Veterinary Association:
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Finally, PUPscan has posted on Facebook a "full report" that will apparently feature on the front page of  Our Dogs magazine. It refers to a "Facebook attack" on PUPscan and efforts to "undermine pedigree dogs and to sabbotage any advancement towards the health and welfare of pedigree dogs by dedicated breeders and professionals". The statement that followed was long and you can read it on their Facebook page or Our Dogs. It doesn't address any of the questions I asked.
​
So, here is where we are.

We don't know what they are measuring. We don't know if they have any evidence that these mystery measurements tell us anything about hip dysplasia. We don't know how measurements of a structure that is cartilage in a puppy can tell us something useful about what to expect in the adult dog after it has been converted to bone. We don't know why they think they can ascribe to genetics any problems they see in their ultrasound examination. 
As far as I can tell, they have no data that link whatever they are measuring to a diagnosis of or predisposition to hip dysplasia. If that's the case, then this is essentially a research project (and note that they call it the "PUPscan Project") in which the owners of the dogs will pay for collection of data that may or may not be useful, and at best it will be several years before they will even be able to say. 

What I find especially disturbing is the fact that they are leading people to believe that they are providing useful information and "new hope for breeders of 'dysplastic' dogs", as in the title of their published article. Unless they can provide answers to the very basic questions I have asked them, I don't see that they have anything useful to offer. 
If you contemplate having them scan your dog, you should ask  them the questions I post at the top. They will need to provide numbers, not verbiage. They will need to provide a published, peer-reviewed report from a respectable journal. Until they can do this, I concur with the the Kennel Club and BVA: continue to participate in the existing screening programs, educate yourself about the relative roles of genetics and environment in expression of hip dysplasia, and take advantage of EBVs offered by various kennel clubs.

You can read the exchange I had with them here:
pupscan_-_timeline.pdf
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Some additional comments on minor issues:
Usefulness of EBVs (Estimated Breeding Values)
PUPscan states that "the EBV method is not yet able to demonstrate genetic abnormalities". I'm not sure what they mean by this; the statement really doesn't make sense. EBVs don't (can't) "demonstrate" genetic abnormalities. They are statistical (hence "estimated") "breeding values", or predictions of the genotype of an animal based on the phenotypes (in this case, hip scores) of related dogs. The EBV score for a dog is based on the calculated heritability of the trait in that population of animals, where heritability is the amount of variation in the trait that can be accounted for by genetic differences among individuals. (For a quick review of what heritability means, check out this nice video from the Kahn Academy.) The heritability of hip dysplasia has been determined in many studies and usually ranges from about 0.15-0.45, which means that 15-45% of the variation among animals reflects variation in genotype; i.e., there is an influence of genetics on hip dysplasia. The EBV score does not "demonstrate genetic abnormalities"; it simply reflects an influence of genotype on phenotype. 

Hip screening over the last 30 years has "not reduced the incidence of Dysplasia"
Certainly hip dysplasia has not been eliminated, but there is evidence that progress has been, or could be, made as a result of screening programs. The Finnish Kennel Club has documented significant improvement in hip phenotype in a number of breeds over the last three decades as a result of traditional screening programs and more recently through the use of Estimated Breeding Values. Similarly, hip evaluations of Hovawarts in Germany were dramatically improved after adoption of a breeding program based on EBVs (http://bit.ly/2l5qReo). Organizations with breeding programs for guide dogs have also had great success in reducing the incidence of hip dysplasia in the animals they produce (pers. comm.). Hip dysplasia continues to be a significant health and welfare issue in dogs, but it is not the case that no progress has been made over the last 30 years. 
NOTE added 1 March 2017

I have posted a followup to this in light of new information that came to my attention.

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Latest OFA statistics for hip dysplasia (Dec 2016)

2/5/2017

 
By Carol Beuchat PhD
The Orthopedic Foundation for Animals (OFA) has updated their website to include the most recent statistics for incidence of hip dysplasia in dogs (through December 2016). The data are reported as number (or %) of dogs with excellent hips and number (or %) of dogs judged to be dysplastic. These data are divided into dogs born in 2011-2015 (so all dogs evaluated are at least 1 year old), and the same statistics for all dogs in the database (including dogs born 2011-2015).
I have downloaded these data and graphed them various ways to make the information more accessible. Because there is a large number of breeds, breed names are almost impossible to read on the whole graphs, but you can download a copy of the file or I have split the graph in two to improve legibility. The data are presented both alphabetically (so you can find the breed you're looking for) and by rank (so you can compare with other breeds). The files for the large graphs can be downloaded.


​There are a few notable points (for dogs born 2011-2015):
  • Best hips: Lhasa Apso & Italian Greyhond
  • Worse hips: Pug & Fila Brasiliero
  • Most improved: Sussex Spaniel & Basset Hound
  • Least improved: Fila Brasiliero & Welsh Terrier
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__excellent__alpha_.png
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excellent_alpha_split.png
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__excellent__rank_.png
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​The graph just above has been split below to make the names a bit easier to read.
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__dysplastic__alpha_.png
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​The graph just above has been split below to make the names a bit easier to read.
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Total & Recent Dysplastic

Indicated for each breed are the % Dysplastic reported in December 2016 for dogs born 2011-2015 (green bar) and the Total % Dysplastic over all years (red bar). If a breed has recent improvement, the green bar will be shorter than the red bar. Only breeds with % Dysplastic < 10% are graphed.
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Same data as above plotted as % Total Dysplasic - % Recent Dysplastic. Bars above the zero line are for breeds that have reduced the incidence of dysplasia; bars below the line are breeds for which recent % Dysplastic is greater (i.e., worse) than the average over all years.
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If hip or elbow dysplasia are problems in your breed,
​check out the great information in ICB's online course!

ICB's course

​Understanding Hip & Elbow Dysplasia
starts 6 February 2017

Sort through the Truth & Myths:

- Is hip dysplasia genetic? Environment? Both?
- Can small dogs have hip dysplasia?
- Will Vitamin C help?
- Is fast growth a risk factor?
- Is exercise good or bad for puppies?
- Does a raw diet prevent hip dysplasia?
- Could it be caused by not enough calcium?
- Can a dog with bad hips produce a puppy with good hips?
- Are bad elbows inherited?
- Is there a DNA test for hip dysplasia?
- Are puppies born with hip dysplasia?
- Do slippery floors cause hip dysplasia?
We'll cover anatomy, biomechanics, nutrition, and the latest research, as well as how you can reduce risk by understanding heritability and estimated breeding values.

The course is online, available anywhere in the world.
Materials are available 24/7
Online course Starts 6 February 2017!

For info and registration, go to - 
http://bit.ly/2iEEh1e

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Why didn't Antarctic sled dogs have hip dysplasia? 

2/2/2017

 
By Carol Beuchat PhD
While digging around for information about the history of sled dogs for my course Biology of Dogs, I ran into a fascinating paper about osteoarthritis in the working dogs that were kept at the scientific field stations on Antarctica for several decades. The thing that was so remarkable is that these dogs developed osteoarthritis by the time they were about 8 years old, but they had no evidence of hip dysplasia. None.

Read on...

In the 1940s, the British established scientific stations on Antarctica. Early on, the program was known as the Falkland Islands Dependencies Survey (FIDS), and it was renamed in 1962 as the British Antarctic Survey.  ​
​For transportation and hauling materials, residents relied on dogs that were bred at the stations from the descendants of the early dogs brought to the continent.
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"The canine population of the Falkland Islands Dependencies is divided between the stations and has fluctuated in number between forty and 200. Dogs are frequently moved from station to station and must be considered as one and not separate breeding units.

There have been three introductions of Huskies into the Falkland Islands Dependencies. Two shipments from Labrador arrived at the beginning of I945 and 1946.  In 1945 about forty dogs came from the coast south of Hopedale and in 1946 another forty from north of this settlement. These animals  were not the best in the area, but rather the small and unwanted beasts. 

In 1954, twenty-one dogs were shipped south; eleven of these had been selected as the best of the British North Greenland Expedition, 1952-54, pack; four more were their progeny; three dogs came from the Northwest Territories of Canada and three from Britain. The last trio were out of a Husky of Labrador parentage and the sire of one pup, although born in Labrador, had won fame in Graham Land as leader to V. E. Fuchs." (Taylor: The breeding and maintenance of sledge dogs)
Because these dogs were essential for both work and survival, the scientists had an interest in learning about their physiology and performance in the harsh conditions. They kept records of both general husbandry and a variety of experiments.

One thing of great interest to them was the working life of the dogs. Most of the dogs were able to pull sleds until about 8 years old, after which arthritis usually reduced performance and the dog was removed. 
"Dog team drivers of the British Antarctic Survey have come to accept the fact that otherwise healthy Antarctic sledge dogs are usually incapable of further work by the time that they are 8 years old. Since this is less than might be expected of a dog of the husky's size and weight under normal conditions, an investigation into the possible causes of the decline of the ageing husky was started in 1963- 64." (Bellars & Godsal 1969)
They evaluated the hips and elbows of 34 dogs that ranged in age from 1.5 to 10 years old. They found no evidence of hip dysplasia in any of them. What they did observe instead were signs of wear and tear associated with hauling loads. 
The erosions of the articular canilage were found to be in a uniform position on the articular surfaces of the humerus and femur. They varied in size and depth, but were invariably bilaterally similar. On the surface of an affected humerus the erosion was on the caudal curvature of the articular cartilage; on the femur it was found on the dorsal aspect, extending cranially and caudally in the more advanced cases. Severe cases also showed smaller erosions on the opposite surfaces of the glenoid cavity of the scapula and the acetabulum. These erosions are found, therefore, in the areas of greatest pressure when a dog is pulling, i.e. on the areas in direct opposition when a dog is leaning against the weight of the loaded sledge, with its limbs in extension retraction and, in the case of the hindlimb, slightly abducted. 
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​Study at post-mortem examination and later showed that there was no evidence of shallowness of the acetabulum in any of the dogs examined. Likewise, the heads of femora and humeri showed normal conformation. There was some re-shaping in the most severely affected cases...The hip joints showed no evidence of sufficient abnormality to be called hip dysplasia. (Bellars & Godsal 1969)
Read that again:

​"The hip joints showed no evidence of sufficient abnormality to be called hip dysplasia."
In the x-rays below, they noted that the hips of the the 1.5 year old dog on the left were completely normal, and the 5 year old dog on the right had evidence of erosion on the head of the femur (at the arrows). The shape and depth of the acetabulum (the socket) in both dogs was normal, as it was in all of the dogs they examined.
​
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They concluded that the decline in working ability of the dogs with age was the result of osteoarthritis of the hip and shoulder joints, and that "hip dysplasia does not occur in a representative selection of these dogs". They further remarked that in the 700 dogs which had been reared at the station, none showed the signs of early hip dysplasia (pain, femoral head subluxation, or gait abnormalities) that are often seen in Labrador Retrievers or German Shepherd Dogs. 
"By virtue of these findings, and the conformation shown in the radiographs, we feel justified in believing that hip dysplasia, as seen in dogs in the United Kingdom, is not present in the Survey's sledge dogs. Such degenerative changes as are apparent are considered to be secondary changes due to aging, exacerbated by "special occupational activities" and "abnormalities of gait"."

​Remember, there was nothing special about the original breeding stock of these dogs. They came from various places, and it was even noted that "These animals were not the best in the area, but rather the small and unwanted beasts.' There were no fancy living accommodations, and dogs had to earn their keep as working sled dogs or they were eliminated from the colony. A portable x-ray machine was brought in to make the films used in the study, but otherwise the dogs were not evaluated for hip structure. Because none of the dogs had hip dysplasia, there was no control program like the ones run today. And remember, these were not small dogs, and they were expected to pull at least their own body weight for dozens or even hundreds of miles, day after day after day. For them, the osteoarthritis that eventually developed was a consequence of aging and biomechanical wear and tear on the articulating surfaces of the femur and humerus.
Many studies of hip dysplasia show it to be a consequence of both genetic and environmental factors, some of which have been studied. But we still don't really know what causes it or how to prevent it. The Antarctic researchers were aware of the high incidence of hip dysplasia among pet dogs in the UK, but they offered no explanation for why their working dogs should be completely free of any evidence of dysplasia. 

​Still, we have more questions than answers.

You can download a copy of the publication ​HERE.

In 1993, the last descendants of the Antarctic sled dogs were ordered to be removed from Antarctica by the Madrid Protocol. This video is about the last trip taken by these iconic dogs and their removal from Antarctica, and it includes some interesting historic footage and information about the history of the Inuit dogs of the Arctic.
​




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​Understanding Hip & Elbow Dysplasia
starts 6 February 2017 

REFERENCES
Bellars ARM & MF Godsal. 1969. Veterinary studies on the British Antarctic Survey's sledge dogs: II. occupational osteoarthritis. British Antarctic Survey Bulletin 22: 15-38.

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